e1b1a in the levantdavid w carter high school yearbook
The Indo-European migrations would certainly have brought some E-V13 early on, from circa 2500 BCE. Z830, M310.1's . CTS1096 split into three subclades around 7,500 to 7,000 years ago, a period that corresponds to the advent of the Copper Age around modern Kurdistan. The Wright Brothers, the inventors of the world's first successful airplane, belonged to haplogroup E-V13 (S7461 subclade). (2012) recovered the DNA of Napoleon Bonaparte from beard hair follicules and compared his Y-DNA to that of one of his present-day descendants, Charles Napolon. Analysis of diversity and rough estimates of times to the most recent common ancestors of haplogroups provide evidence of multiple expansions along eastern and western routes and a late, exclusively eastern route, expansion. E1B1A must be the standard for determining whether or not a male is a descendant of the Biblical Israelites. The increase in the rate of identification of slowly mutating NRY binary markers (ie, unique event polymorphisms (UEPs))21, 22, 23 has resulted in many studies designed to investigate the paternally mediated genetic relationships of sub-Saharan African populations. E1b1a is an African lineage that expanded from northern Africa to sub-Saharan and equatorial Africa with the Bantu agricultural expansion. The genetic data are thus in broad agreement with analysis based on linguistic studies, which suggests that the spread of Bantu languages is the consequence of successive dispersals and that a single large-scale migration by Bantu speakers is unlikely.3 It is also consistent with suggestions that differences between eastern and western Bantu languages are a consequence of expansion patterns.3 This interpretation suggests the absence of substantial male-mediated gene flow from East-Central Africa to West-Central Africa during the past millennium, because had it occurred, it would be expected that examples of haplogroup E1b1a8a1a would have been observed in the Congolese groups included in this study. The original Phoenician M81 in the Levant could also have diffused across the Eastern Mediterranean over the centuries, during the Roman, Byzantine and Ottoman periods. Google Scholar. Nei M : Molecular Evolutionary Genetics. Sectors in pie charts are coloured according to the haplogroup colour code to the left. [6][7][8][9] According to Wood et al. Genome Res 2008; 18: 830838. . John's father, Matthias Corvinus (1443-1490) was King of Hungary and Croatia, and disputed King of Bohemia and Duke of Austria. do you know whether the hp E1b1a was ever found in ancient Levant? There are at least three distinct sources of E-V13 in Italy. Indeed the distribution pattern and frequency of M81 matches much better the Phoenician maritime empire, with its origins in the Levant, and its dispersal along the cost of North Africa, but also Iberia, Sardinia and Sicily. One of them was E-M34 (notably Levantine clades like Y15558 and Z21421), which makes up about 15% of modern Lebanese Y-DNA, but was probably higher before the Greek, Roman, Arabic, Byzantine, medieval crusader and Ottoman occupations altered the local gene pool. [30] E-M10 was found in a single person of the Lissongo group in the Central African Republic and two members in a "Mixed" population from the Adamawa region.[12]. Haplogroup E-M2, also known as E1b1a1-M2, is a human Y-chromosome DNA haplogroup. E1b1a is also known as E-M2 and E1b1b is also know as E-M215 or as E-M35. L2a is widespread in Africa and the most common and . Iranic tribes, La Tne Celts, Romans, Goths, Slavs). The discovery of two SNPs (V38 and V100) by Trombetta et al. Cruciani et al. Y-DNA Haplogroup E: E1b1b and E1b1a - Your DNA Guide Did E1b1a mutate from E1b1b? And if it did - ProBoards As both NRY and mtDNA genetic systems have smaller effective population sizes than autosomal markers, they are more prone to genetic drift14, 15, 16 and are therefore more likely to differ among groups than are autosomal markers. [28][27] The ancestral sickle cell haplotype to modern haplotypes (e.g., Cameroon/Central African Republic and Benin/Senegal haplotypes) may have first arose in the ancestors of modern West Africans, bearing haplogroups E1b1a1-L485 and E1b1a1-U175 or their ancestral haplogroup E1b1a1-M4732. Lazaridis et al. The American actor and producer Nicolas Cage (born 1964),has been found to belong to haplogroup E1b1b-M84. The ancestral L485 SNP (along with several of its subclades) was very recently discovered. So I was wondering if such a marker has anything to do with the Natufian Neolithic culture of the Levant as some of the skulls associated with this particular culture have been described as Sub-Saharan-like. E1b1a (L576) This population represents an East to West thrust in Africa, only E1b1a lineage able to survive crossing the A1b1 territories. This includes some E1b1b subclades like V22 (12,000 years old) and V32 (10,000 years old), but also undeniably Near Eastern lineages like T1a-CTS2214 and J1-L136. However, since G2a is the only lineage that was consistently found in all Neolithic sites tested to date in Europe, the absence of Neolithic G2a lineages from Scandinavia and the Baltic implies that no Neolithic lineage survives there, and consequently E-V13 does not date from the Neolithic in the region. Giuseppe Garibaldi (1807-1882), the general, politician and nationalist who played a large role in the history of Italy, probably belonged to haplogroup E-V13 based on the Y-DNA results from another Garibaldi from the same province in his ancestral Liguria. The PF6759 subclade seems to have reached Sardinia during the Neolithic period. The basal node E-L485* appears to be somewhat uncommon but has not been sufficiently tested in large populations. why EPF2431 is rare - eupedia.com Wairak people in Tanzania tested 4.6% (2/43) positive for E-M10. mtDNA variability in two Bantu-speaking populations (Shona and Hutu) from Eastern Africa: implications for peopling and migration patterns in sub-Saharan Africa. All of the groups characterised in this study speak a Niger-Congo language, except for the Anuak in south-west Ethiopia who speak a Nilo-Saharan language. The haplogroup E1b1a-M2 (and its sub-lineages) is widely spread in Africa and highly prevalent in all Bantu sub-Saharan populations, with frequencies above 80% in most populations 39,40,46,47. Tanya M Simms 2011, The Peopling of the Bahamas: A Phylogeographical the migration of a small group of settlers carrying among whom one paternal lineage was much more common than any others. Living Descendants of Biblical Canaanites Identified Via DNA According to the DNA results of a relative, Google co-founder Larry Page (b. Tishkoff SA, Reed FA, Friedlaender FR et al. [33] In other words, as one moves to West Africa from western Central Africa, the less subclade E1b1a1f is found. Edmonds CA, Lillie AS, Cavalli-Sforza LL : Mutations arising in the wave front of an expanding population. In Anatolia, E-V13 is found mostly in the western third of the country, the region that used to belong to ancient Greece. The Genomic History of the Bronze Age Southern Levant [26] West Africans (e.g., Mende of Sierra Leone), bearing the Senegal sickle cell haplotype,[29][26] may have migrated into Mauritania (77% modern rate of occurrence) and Senegal (100%); they may also have migrated across the Sahara, into North Africa, and from North Africa, into Southern Europe, Turkey, and a region near northern Iraq and southern Turkey. For many years the vast majority of academics have assumed that E-V13 and other E1b1b lineages came to the Balkans from the southern Levant via Anatolia during the Neolithic, and that the high frequency of E-V13 was caused by a founder effect among the colonisers. Traces of earlier inhabitants, however, can be observed today in these regions via the presence of the Y DNA haplogroups A1a, A1b, A2, A3, and B-M60 that are common in certain populations, such as the Mbuti and Khoisan. Multiple origins of Ashkenazi Levites:Y chromosome evidence for both Near Eastern and European ancestries. [20], At Cabeo da Amoreira, in Portugal, an enslaved West African man, who may have been from the Senegambian coastal region of Gambia, Mauritania, or Senegal, and carried haplogroups E1b1a and L3b1a, was buried among shell middens between the 16th century CE and the 18th century CE. Castri L, Tofanelli S, Garagnani P et al. [25] Pita was of Sub-Saharan African ancestry and carried haplogroups E1b1a-M4287 and L3e2b. Nowadays E-M81 is the dominant paternal lineage among Northwest Africans, and particularly Tuaregs, Mountain Moroccans, Tunisians and Libyans. The small presence of E-V13 in the Near East could be better explained by the extremely long Greek presence in the eastern Mediterranean from the time of Alexander the Great until the end of the Byzantine domination over the region during the Middle Ages. Underhill PA, Jin L, Lin AA et al. [5] The downstream SNP E-M180 may have originated in the humid south-central Saharan savanna/grassland of North Africa between 14,000 BP and 10,000 BP. It is believed to have first appeared in the Horn of Africa approximately 26,000 years ago and dispersed to North Africa and the Near East during the late Paleolithic and Mesolithic periods. Mitochondrial, Y-chromosome and autosomal DNA analyses have been carried out in attempts to understand the demographic events that have taken place. Montano et al. For other uses, see. Haplogroup E-V68 - Wikipedia Distribution of haplogroup E-M123 in Europe, the Middle East & North Africa. https://doi.org/10.1038/ejhg.2012.176, DOI: https://doi.org/10.1038/ejhg.2012.176. E-M34 is the main Middle Eastern variety of E1b1b and is thought to have arrived with the Proto-Semitic people in the Late Copper to Early Bronze Age. The distribution of haplogroup E1b1a8a1* defined by U290 in the absence of U181 with a TMRCA of 14131725 YBP is similar to that of E1b1a8 and may be interpreted in the same way. As of November 2016, he was the 12th richest person in the world. Searching for the roots of the first free African American - Nature Franz Kafka, a German-speaking Bohemian novelist and short-story writer, who is widely regarded as one of the major figures of 20th-century literature probably belonged to E-Y161794, a Jewish branch of haplogroup E-M81, based on the Y-DNA test of another Kafka from Czechia at FTDNA. Because of the separation of the two major waves around 3500 YBP and the subsequent isolation of many groups, modern Bantu languages can be divided into West and East Bantu.8, 9 Non-Bantu languages (that do not belong to the Niger-Congo phylum, eg, Khoisanid languages in the southwest of the continent and a few Cushitic and Nilo-Saharan tongues in the northeast6) have survived in areas that have not experienced extensive inward migration of Bantu speakers. This indicates that a single man may have had nine sons who went on to have numerous children of their own. As a consequence, this study makes an important contribution to filling the gap. Although sampling in most NRY studies of sub-Saharan Africa has, in the past, been quite limited in terms of geographic coverage and sample sizes, the distribution of this haplogroup is relatively well described in groups living along both the postulated western and eastern routes of the EBSP, as well as in Senegal29 and Cameroon27, 30 in West Africa. In this study, we analyse, as did Alves et al,33 both UEP and short tandem repeat (STR) (in this study restricted to NRY) to show that geographic frequency distributions and the time to the most recent common ancestors (TMRCAs) of haplogroups, comprising haplogroup E1b1a in 43 sub-Saharan African groups (n=2757) with diverse linguistic affiliations (Supplementary Figure S1), reveal multiple waves of expansion from West Africa, with a late expansion along the eastern route but not the western. [2] E-M329 is also frequent in Southwestern Ethiopia, especially among Omotic -speaking populations. Also downstream of CTS1096, the Y14891 and Z21018 clades are typically found among people of Jewish ancestry, while PF6391 and Z21421 are found in the Levant (Syria, Lebanon, Palestine, Jordan) and the Arabian peninsula. It would be easy to assume that E-M81 colonised Northwest Africa during the Mesolithic or Neolithic period, then spread to southern and western Europe with the southern wave of Neolithic farmers that crossed over from Morocco to Iberia, then spread around western Europe with the Megalithic people. Veeramah et al. Z830, M310.1's brother clade, is almost exclusively Middle Eastern. Genetic history of the Middle East - Wikipedia Although it is generally accepted that the EBSP has its origin in the so-called Bantu Homeland situated in the area of the border between Nigeria and the Grassfields of Cameroon, and that it followed both western and eastern routes, much less is known about the number and dates of those expansions, if more than one. E1b1b used to be E3b, but always is E-M215 or E-M35. Behar DM, Thomas MG, Skorecki K et al. According to the equation, the minimum frequency at which a haplotype is present for it to have a 95% probability of being observed, given that n chromosomes are typed, is q=110(log(0.05)/n). [29] West Africans, bearing the Benin sickle cell haplotype, may have migrated into the northern region of Iraq (69.5%), Jordan (80%), Lebanon (73%), Oman (52.1%), and Egypt (80.8%). L2 has five main subhaplogroups: L2a, L2b, L2c, L2d and L2e. PubMed L791 and Z21466 have a mostly European distribution today and their ages point toward a Neolithic diffusion. New Jersey: Princeton University Press, 1994. [69] This is the modal haplotype of STR markers that is common in carriers of E-U175. Haplogroup E1b1a7 (defined by M191) is modal in most groups in countries from Ghana to Mozambique and only at slightly lower frequency in South African Bantu speakers (33.8% compared with E1b1a8*. Haplogroup L2 (mtDNA) - Wikipedia E-U175 and E-L485) of E1b1a evolved. Therefore both hypotheses are plausible. See also : Southern Neolithic route brought Megaliths from the Levant to Western Europe. It would be unthinkable that over 1,500 years of Hellenisation and Byzantine rule in Anatolia and the Levant didn't leave any genetic trace. Some of these SNPs have little or no published population data and/or have yet to receive nomenclature recognition by the YCC. It's typical of all E1b1b haplogroups, but E1b1a has instead 438=11 and only 2% of E1b1a samples have 438=10. Because the West-Central African E1b1a data set is sufficiently large (n=516; eight groups), we would have expected to observe the E1b1a8a1a haplotype, if present at a frequency as low as 0.0058. Weale ME, Shah T, Jones AL et al. found similarly low frequencies of basal E-U175* in subjects in the Ivory Coast and Benin. [25] Tima was of western Central African ancestry and carried haplogroup L3e1e. Grard Lucotte et al. In just a few centuries, that very minor E-V13 lineage had started an expansion process that would turn it into one of Europe's most widespread paternal lineages and reach far beyond the borders of Europe itself, also spreading to the eastern edge of the Mediterranean, the Caucasus, Kurdistan, Iran, and even Siberia. The African diaspora: mitochondrial DNA and the Atlantic slave trade. They would have brought typically Germanic lineages like I1 and R1b-U106, but also the Proto-Slavic R1a-CTS1211, which is now found uniformly in 1 to 2% of the population. Haplogroup E-V38 - Wikipedia and JavaScript. Whilst E1b1a reaches its highest frequency of 81% in Senegal, only 1 of the 139 Senegalese that were tested showed M191/P86. Am J Hum Genet 2004; 74: 532544. There is evidence that the Natufians already cultivated cereals like rye before the Neolithic period. This page has been accessed 678 times. Am J Hum Genet 2000; 66: 674686. "E3a" redirects here. Table 1 reports the frequencies of all observed haplogroups, including the component haplogroups of E1b1a. You should learn them by the mutations because the letters change, the mutations don't. E1b1a used to be E3a, but always was E-M2. We analyse frequencies of halpogroups and estimates of TMRCA to answer two questions: (a) Is there evidence of more than one expansion of paternal line ancestors of Bantu-speaking people living in present day sub-Saharan Africa? The authors declare no conflict of interest. The probability of observing a particular haplotype, if present, in a randomly collected set was assessed by the equation (1q)n=(1P), where P is the probability of observing the haplotype, q is the minimum frequency of the haplotype to be observed and n is the number of chromosomes. Variation of female and male lineages in sub-Saharan populations: the importance of sociocultural factors. Last update February 2023 (famous members). Consequently, the haplogroup is often observed in the United States populations in men who self-identify as African Americans. They established that both men belonged to haplogroup E-M34, a subclade which is thought to have reached Mediterranean Europe from the Levant during the Neolithic period. Searching for the roots of the first free African American community, Carriers of mitochondrial DNA macrohaplogroup L3 basal lineages migrated back to Africa from Asia around 70,000 years ago, The peopling of the last Green Sahara revealed by high-coverage resequencing of trans-Saharan patrilineages. Y6923 also emerged around 3500 BCE, but became almost extinct. Evidence from Y-chromosome analysis for a late exclusively eastern The J haplogroup is of Semitic origin and is overwhelmingly present in The Middle East. [69], The supposed "Bantu haplotype" found in E-U175 carriers is "present at appreciable frequencies in other NigerCongo languages speaking peoples as far west as Guinea-Bissau". However, Razib Khan in this podcast says that E1b1a was pretty common among ancient Levantines. There is clearly a radiation from the Greece (where E-V13 makes up approximately 30% of the paternal lineages) to the East Mediterranean (where the frequency drops to under 5%). Salas A, Richards M, De la FT et al. Subsequently, the expansion is thought to have continued along the south-eastern coast (East-Bantu route).5 In an alternative model, the split came later after passage through the rain forest.3, 4 The Bantu language family is distributed throughout most of sub-equatorial Africa and is the continents largest, both in terms of the numbers of individuals speaking it and its geographic spread.2, 6, 7 This level of linguistic homogeneity among geographically distant populations across sub-Saharan Africa supports the suggestion of rapid expansion. E1b1a1a1 is commonly defined by M180/P88. Combined use of biallelic and microsatellite Y-chromosome polymorphisms to infer affinities among African populations. Furthermore, all the modern members of E-V13 descend from a common ancestor who lived approximately 5,500 years ago, and all of them also descend from a later common ancestor who carried the CTS5856 mutation. [17][18], At a San Jose de los Naturales Royal Hospital burial site, in Mexico City, Mexico, three enslaved West Africans of West African and Southern African ancestry, dated between 1453 CE and 1626 CE, 1450 CE and 1620 CE, and 1436 CE and 1472 CE, were found; one carried haplogroups E1b1a1a1c1b/E-M263.2 and L1b2a, another carried haplogroups E1b1a1a1d1/E-P278.1/E-M425 and L3d1a1a, and the last carried haplogroups E1b1a1a1c1a1c/E-CTS8030 and L3e1a1a. The material culture of the Late Chalcolithic period in the southern Levant (4500-3900/3800 BCE) is qualitatively distinct from previous and subsequent periods. The Phoenicians would have spread E-M34 to Cyprus, Malta, Sicily, Sardinia, Ibiza and southern Iberia. ISSN 1018-4813 (print), Evidence from Y-chromosome analysis for a late exclusively eastern expansion of the Bantu-speaking people, Subdividing Y-chromosome haplogroup R1a1 reveals Norse Viking dispersal lineages in Britain, The role of matrilineality in shaping patterns of Y chromosome and mtDNA sequence variation in southwestern Angola, Autosomal genetics and Y-chromosome haplogroup L1b-M317 reveal Mount Lebanon Maronites as a persistently non-emigrating population, Y-chromosomal connection between Hungarians and geographically distant populations of the Ural Mountain region and West Siberia, A comprehensive portrait of Y-STR diversity of Indian populations and comparison with 129 worldwide populations, Origin and diffusion of human Y chromosome haplogroup J1-M267, The paternal and maternal genetic history of Vietnamese populations, North Asian population relationships in a global context, Early medieval genetic data from Ural region evaluated in the light of archaeological evidence of ancient Hungarians, Supplementary Tables S1-S2-S4 (DOC 141 kb), Distribution of paternal lineages in Mestizo populations throughout Mexico: an in silico study based on Y-STR haplotypes. These data are consistent with multiple expansion events southwards from West Africa. The Phoenicians possessed a variety of paternal lineages reflecting the complex ancient history of the Middle East. volume21,pages 423429 (2013)Cite this article. Anthropology, archaeology, linguistics and, in recent decades, genetics have been used to elucidate some of the events and processes involved. For comparison, the NRY haplotype diversity treating E1b1a as a single haplogroup ranged from 0.821 to 0.945, with the exception of Anuak who displayed a much lower diversity (h=0.516). It was first reported in a person from the Gambia.[76]. Nevertheless, many lineages now found among the Ethiopians and Somalians appear to have come from the Fertile Crescent during the Neolithic period. A combination of the two scenarios could provide an even better explanation. E1b1a1a1b is defined by M116.2, a private marker. E-M78 and E-Z827 originated respectively at 20,000 years and 24,000 years. (2007) suggests that E-M78, E1b1b predominant subclade in Egypt, originated in "Northeastern Africa", with a corridor for bidirectional migrations between northeastern and eastern Africa (at least 2 episodes between 23.9-17.3 ky and 18.0-5.9 ky ago), trans-Mediterranean migrations directly from northern Africa to Europe (mainly in [12], E1b1a1a1e is defined by markers M10, M66, M156 and M195. At present the most consistent explanation is that E-V13 developed from E-M78 in Central or Eastern Europe during the Neolithic period, and was assimilated by the R1a and R1b Proto-Indo-Europeans around the time that they were leaving the Pontic Steppe to invade the rest of Europe. In . The earliest known prehistoric sample to date is an E-V13 from Catalonia dating from 5000 BCE. New Haven: Yale University Press, 1995. (2011) only found one out of 505 tested African subjects who was U175 positive but negative for U209. [25] Lisa was of West African ancestry and carried haplogroups E1b1a-Z6020 and H100. [25], Amid the Green Sahara, the mutation for sickle cell originated in the Sahara[26] or in the northwest forest region of western Central Africa (e.g., Cameroon)[26][27] by at least 7,300 years ago,[26][27] though possibly as early as 22,000 years ago. remains uncertain. 2018). The study revealed that he belonged to haplogroup E1b1b1. The absence of E-V13 from Central Anatolia does not concord with a diffusion linked to Neolithic agriculture. peoples). All modern carriers of this lineage descend from a common ancestor who lived only 1,200 years ago, and all are Ashkenazi Jews. Of course, the TMRCA is only an estimate and could vary by a few centuries.
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